| Original Botanical Description or Link to | Perennial, monoecious herb, semi-erect or appressed to substrate at the base and distally erect, to c. 30 cm tall. Stem branched; internodes 2–6 cm long, brownish-reddish, with microscopic glandular hairs and a sparse indumentum of multicellular red hairs, c. 0.5−1.5 mm long. Stipules caducous, 8–12 × 4–5.5 mm, elliptic, slightly anisophyllus, with an abaxially slightly prominent midrib, apex narrowed into a bristle projecting
up to 1.5 mm, reddish greenish, translucent at the margin, abaxially glabrous. Leaves: petioles 2–6 cm long, adaxially shallowly channelled, reddish, moderately covered with red hairs, denser on the younger parts, c. 1.5 mm long; lamina 9.5−14 × 7−9.5 mm, asymmetric, ovate to elliptic, base cordate and lobes not or just slightly overlapping, apex acuminate, margin denticulate to serrulate, adaxial surface green-reddish, with
reddish veins toward the base and pale green towards the margin, glabrous, abaxially pale green-reddish, sparsely hairy on the veins only; venation palmate-pinnate, primary veins 6–7, actinodromous, secondary veins craspedodromous. Inflorescences protogynous; female inflorescences 2-flowered, sometimes associated with a single male flower in between, or single female flower with single male flower, peduncles
c. 5 mm long, glabrescent to glabrous; male inflorescences paniculate-cymose with 3−4 dichasial-monochasial partial inflorescences on up to 2 cm long peduncles in the basal part, each with up to 8 flowers, bracts caducous. Male flowers: pedicels 1.5–3.5 cm long, white, greenish or reddish, glabrous; tepals 2, white tinged pink, 9–12 × 11–14 mm, broadly ovate, base slightly cordate, apex rounded, outer surface glabrous; androecium of c. 26–27 stamens, yellow, filaments up to c. 0.3–0.8 mm long, slightly fused at the very base, anthers up to c. 0.7–1 mm long, oblong to narrowly obovate,
dehiscing through unilaterally positioned slits that are c. ½ as long as the anthers. Female flowers: pedicels 1–2 cm long, reddish, glabrous; tepals 5, white tinged pink, unequal, one smaller 8–10 × 4–5 mm, elliptic, the four larger 11–12 × 8–10 mm, ovate, outer surface glabrous; ovary (excluding wings) 8–10 × 4.5–5 mm, ellipsoid, pink to red, glabrous, locules 3, placentation axile, placentae bilamellate, wings 3, subequal wing slightly larger than two others to unequal, pale green-reddish, base rounded, apex
subtruncate to truncate, up to 5–7 mm at widest point (apically or subapically); style c. 6 mm long, basally fused, 3-branched, each stylodium bifurcate in the stigmatic region, stigmatic surface a spirally twisted papillose band, orange. Fruits: peduncles c. 5 mm long; pedicels 1.5–2 cm long, pendulous; seed-bearing part ellipsoid, 8–11 × 4.5–5 mm (excluding the wings), glabrous, dehiscent, splitting along the wing attachment, wings as for ovary, up to 8 mm at the widest point (apically or subapically). Seeds
barrel-shaped, c. 0.2–0.6 mm long. |
